Sam Steyaert
Associate Professor
Faculty of Biosciences and Aquaculture
Carrion ecology became my main research focus during recent years. With our REINCAR project, we use various methods (from molecular tools to remote sensing approached) to explore how large vertebrate carrion affects various ecological communities (e.g. vegetation, fungi, scavengers, etc.) and modulates ecological processes (e.g. directed seed dispersal) on both local and landscape scales. We use both experimental and observational approaches and have study areas in the alpine and Arctic tundra, as well as in the boreal forest.
I a teacher and course responsible in the MSc level courses BIO5020 (Landscape ecology and nature management) and GEO5020 (Remote sensing and nature management). Furthermore, I supervise students within the individual based 'special pensum' courses (DR435F/BI308F) in topics such as species distribution modeling and urban ecology, and supervise several BSc, MSc, and PhD students.
2024
From death comes life: Large vertebrate carrion enhances seedling establishment in clonal ericaceous shrubs 2024, Functional Ecology: Arnberg, Mie Prik; Eycott, Amy Elizabeth; Steyaert, Sam
2023
Demographic Parameters of Asian Black Bears in Central Japan 2023, Mammal Study: Volum 48 s. 231-244 Tochigi, Kahoko; Steyaert, Sam; Fukasawa, Keita; Kuroe, Misako; Anezaki, Tomoko; Naganuma, Tomoko
Perfect poopers; passerine birds facilitate sexual reproduction in clonal keystone plants of the boreal forest through directed endozoochory towards dead wood 2023, Forest Ecology and Management: Volum 532 Arnberg, Mie Prik; Patten, Michael A.; Klanderud, Kari; Haddad, Camela; Larsen, Oddbjørn; Steyaert, Sam
Nutrient intake and its possible drivers in free-ranging European brown bears (Ursus arctos arctos) 2023, Ecology and Evolution: Volum 13 De Cuyper, Annelies; Strubbe, Diederik; Clauss, Marcus; Lens, Luc; Zedrosser, Andreas; Steyaert, Sam
The social system of a “nonsocial” species, the brown bear Bear and Human: Facets of a Multi-Layered Relationship From Past to Recent Times with an Emphasis on Northern Europe 2023, Brepols: s. 121-128 Zedrosser, Andreas; Frank, Shane; Hansen, Jennifer Eve; Steyaert, Sam; Swenson, Jon
Gut microbiome biogeography in reindeer supersedes millennia of ecological and evolutionary separation 2023, FEMS Microbiology Ecology: Volum 99 Kamenova, Stefaniya; de Muinck, Eric; Veiberg, Vebjørn; Utsi, Tove Hilde Ågnes; Steyaert, Sam; Albon, Steve
2022
Differentiation and seasonality in suitable microsites of seed dispersal by an assemblage of omnivorous mammals 2022, Global Ecology and Conservation: Volum 40 Tochigi, Kahoko; Steyaert, Sam; Naganuma, Tomoko; Yamazaki, Koji; Koike, Shinsuke
Directed endozoochorous dispersal by scavengers facilitate sexual reproduction in otherwise clonal plants at cadaver sites 2022, Ecology and Evolution: Volum 12 Arnberg, Mie Prik; Frank, Shane; Blaalid, Rakel; Davey, Marie Louise; Eycott, Amy Elizabeth; Steyaert, Sam
Land cover type modulates the distribution of litter in a Nordic cultural landscape 2022, PLOS ONE: Volum 17 Eide, Sara Kristiane Kjærgård; Leh, Linn Nefertari; EInes, Katinka Svatun; Hovland, Ingunn; By, Marit; Ingvaldsen, Elise Wiseth
2021
Grading fecal consistency in an omnivorous carnivore, the brown bear: Abandoning the concept of uniform feces 2021, Zoo Biology: Volum 40 s. 182-191 De Cuyper, Annelies; Clauss, Marcus; Lens, Luc; Strubbe, Diederik; Zedrosser, Andreas; Steyaert, Sam
Size is not everything: differing activity and foraging patterns between the sexes in a monomorphic mammal 2021, Behavioral Ecology and Sociobiology: Volum 75 s. 1-14 Lodberg-Holm, Hanna Kavli; Steyaert, S.M.J.G.; Reinhardt, Stefanie; Rosell, Frank Narve
Animal-borne video systems provide insight into the reproductive behavior of the Asian black bear. 2021, Ecology and Evolution: Volum 11 s. 9182-9190 Naganuma, Tomoko; Tanaka, Mii; Tezuka, Shiori; Steyaert, Sam; Tochigi, Kahoko; Inagaki, Akino
2020
Brown bear (Ursus arctos; Eurasia) | Brown bear (Ursus arctos; Eurasia) | Bears of the World. Ecology, conservation and management 2020, Cambridge University Press: s. 139-161 Swenson, Jon; Ambarli, Hüseyin; Arnemo, Jon Martin; Baskin, Leonid; Ciucci, Paolo; Danilov, Pjotr I.
Mating Strategies Bears of the World. Ecology, conservation and management 2020, Cambridge University Press: s. 21-35 Steyaert, Sam; Zedrosser, Andreas; Swaisgood, R; Filipczyková, Eva; Crudge, Brian Patrick; Dutta, Trishna
Fear the reaper: ungulate carcasses may generate an ephemeral landscape of fear for rodents: Rodents fear scavengers at mass die-off 2020, Royal Society Open Science: Volum 7 s. 1-11 Frank, Shane; Blaalid, Rakel; Mayer, Martin; Zedrosser, Andreas; Steyaert, Sam
Beaver (Castor fiber) activity and spatial movement in response to light and weather conditions 2020, Mammalian Biology: Volum 100 s. 261-271 Bartra Cabré, Laura; Mayer, Martin; Steyaert, Sam; Rosell, Frank Narve
2019
Endozoochory by brown bears stimulates germination in bilberry 2019, Wildlife Biology: Volum 2019 Steyaert, Sam; Hertel, Anne Gabriela; Swenson, Jon
Reindeer carcasses provide foraging habitat for insectivorous birds of the alpine tundra 2019, Ornis Norvegica: Volum 42 s. 36-40 Badia, Rudy; Steyaert, Sam; Puliti, Stefano; Blaalid, Rakel; Frank, Shane
Proximity to humans is associated with longer maternal care in brown bears 2019, Behavioral Ecology and Sociobiology: Volum 73 Van de Walle, Joanie; Leclerc, Martin; Steyaert, Sam; Zedrosser, Andreas; Swenson, Jon; Pelletier, Fanie
A human-induced landscape of fear influences foraging behavior of brown bears 2019, Basic and Applied Ecology: Volum 35 s. 18-27 Lodberg-Holm, Hanna Kavli; Gelink, Henriette Wathne; Hertel, Anne Gabriela; Swenson, Jon; Domevscik, Matej; Steyaert, Sam
Heritability of head size in a hunted large carnivore, the brown bear (Ursus arctos) | Heritability of head size in a hunted large carnivore, the brown bear (Ursus arctos) | 2019, Evolutionary Applications: Volum 12 s. 1124-1135 Rivrud, Inger Maren; Frank, Shane; Bischof, Richard; Mysterud, Atle; Steyaert, Sam; Hertel, Anne Gabriela
2018
Diversionary feeding can reduce red deer habitat selection pressure on vulnerable forest stands, but is not a panacea for red deer damage | Diversionary feeding can reduce red deer habitat selection pressure on vulnerable forest stands, but is not a panacea for red deer damage | 2018, Forest Ecology and Management: Volum 407 s. 166-173 Arnold, Johanna Maria; Gerhardt, Philipp; Steyaert, Sam M.J.G.; Hochbichler, Eduard; Hackländer, Klaus
Special delivery: scavengers direct seed dispersal towards ungulate carcasses 2018, Biology Letters: Volum 14 Steyaert, Sam; Frank, Shane; Puliti, Stefano; Badia, Rudy; Arnberg, Mie P.; Beardsley, Jack
Rub-tree selection by Andean bears in the Peruvian dry forest 2018, Ursus (International Association for Bear Research and Management): Volum 29 s. 58-66 Kleiner, Jack D.; Horn, Russell C. Van; Swenson, Jon; Steyaert, Sam
2017
Marking behavior of Andean bears in an Ecuadorian cloud forest: A pilot study 2017, Ursus (International Association for Bear Research and Management): Volum 27 s. 122-128 Filipczyková, Eva; Heitkönig, Ignas M.A.; Castellanos, Armando; Hantson, Wouter; Steyaert, Sam
Hunting promotes sexual conflict in brown bears 2017, Journal of Animal Ecology: Volum 86 s. 35-42 Gosselin, Jacinthe; Leclerc, Martin; Zedrosser, Andreas; Steyaert, Sam M.J.G.; Swenson, Jon; Pelletier, Fanie
Caught in the mesh: roads and their network-scale impediment to animal movement 2017, Ecography: Volum 40 s. 1369-1380 Bischof, Richard; Steyaert, Sam M.J.G.; Kindberg, Jonas
Indirect effects of bear hunting: a review from Scandinavia 2017, Ursus (International Association for Bear Research and Management): Volum 28 s. 150-164 Frank, Shane; Ordiz Fernandez, Andres Avelino; Gosselin, Jacinthe; Hertel, Anne Gabriela; Kindberg, Jonas; Leclerc, Martin
2016
Bears and berries: species-specific selective foraging on a patchily distributed food resource in a human-altered landscape 2016, Behavioral Ecology and Sociobiology: Volum 70 s. 831-842 Hertel, Anne Gabriela; Steyaert, Sam; Zedrosser, Andreas; Mysterud, Atle; Lodberg-Holm, Hanna Kavli; Gelink, Henriette Wathne
Ecological implications from spatial patterns in human-caused brown bear mortality 2016, Wildlife Biology: Volum 22 s. 144-152 Steyaert, Sam; Zedrosser, Andreas; Elfström, Marcus; Ordiz Fernandez, Andres Avelino; Leclerc, Martin; Frank, Shane
Human shields mediate sexual conflict in a top predator 2016, Proceedings of the Royal Society of London. Biological Sciences: Volum 283 Steyaert, Sam; Leclerc, Martin; Pelletier, Fanie; Kindberg, Jonas; Brunberg, Sven; Swenson, Jon
Seasonal and annual variation in the diet of brown bears Ursus arctos in the boreal forest of southcentral Sweden | Seasonal and annual variation in the diet of brown bears Ursus arctos in the boreal forest of southcentral Sweden | 2016, Wildlife Biology: Volum 22 s. 107-116 Stenset, Nina Emilie; Lutnæs, Paul Antoni Nilsen; Bjarnadóttir, Valgerður; Dahle, Bjørn Steinar; Fossum, Kristin Høivik; Jigsved, Patrick
Temporal effects of hunting on foraging behavior of an apex predator: Do bears forego foraging when risk is high? 2016, Oecologia: Volum 182 s. 1019-1029 Hertel, Anne Gabriela; Zedrosser, Andreas; Mysterud, Atle; Støen, Ole-Gunnar; Steyaert, Sam; Swenson, Jon
Reindeer habitat selection under the risk of brown bear predation during the calving season 2016, Ecosphere: Volum 7 Sivertsen, Therese Ramberg; Åhman, Birgitta; Steyaert, Sam M.J.G.; Rönnegård, Lars; Frank, Jens; Segerström, Peter
2015
A "clearcut" case? Brown bear selection of coarse woody debris and carpenter ants on clearcuts 2015, Forest Ecology and Management: Volum 348 s. 164-173 Frank, Shane; Steyaert, Sam; Swenson, Jon; Storch, Ilse; Kindberg, Jonas; Barck, Hanna
Socio‑ecological features other than sex affect habitat selection in the socially obligate monogamous Eurasian beaver 2015, Oecologia: Volum 179 s. 1023-1032 Steyaert, Sam; Zedrosser, Andreas; Rosell, Frank Narve
2014
Litter loss triggers estrus in a nonsocial seasonal breeder 2014, Ecology and Evolution: Volum 4 s. 300-310 Steyaert, Sam M.J.G; Swenson, Jon; Zedrosser, Andreas
Behavioral correlates of supplementary feeding of wildlife: Can general conclusions be drawn? 2014, Basic and Applied Ecology: Volum 15 s. 669-676 Steyaert, Sam M.J.G; Kindberg, Jonas; Jerina, Klemen; Krofel, Miha; Stergar, Matija; Swenson, Jon
2013
Infanticide as a male reproductive strategy has a nutritive risk effect in brown bears 2013, Biology Letters: Volum 9 Steyaert, Sam M.J.G; Reusch, C.; Brunberg, Sven; Swenson, Jon; Hackländer, K.; Zedrosser, Andreas
Male reproductive strategy explains spatiotemporal segregation in brown bears 2013, Journal of Animal Ecology: Volum 82 s. 836-845 Steyaert, Sam M.J.G; Kindberg, Jonas; Swenson, Jon; Zedrosser, Andreas
2012
The mating system of the brown bear Ursus arctos | The mating system of the brown bear Ursus arctos | 2012, Mammal Review: Volum 42 s. 12-34 Steyaert, Sam M.J.G; Endrestøl, Anders; Hackländer, Klaus; Swenson, Jon; Zedrosser, Andreas
Faecal spectroscopy: a practical tool to assess diet quality in an opportunistic omnivore 2012, Wildlife Biology: Volum 18 s. 431-438 Steyaert, Sam M.J.G; Hütter, Franziska J.; Elfström, Marcus; Zedrosser, Andreas; Hackländer, Klaus; Le, Minh Hung
2011
Brown bear conservation and the ghost of persecution past 2011, Biological Conservation: Volum 144 s. 2163-2170 Zedrosser, Andreas; Steyaert, Sam Mathias Jozef Georges; Gossow, Hartmut; Swenson, Jon
Resource selection by sympatric free-ranging dairy cattle and brown bears Ursus arctos | Resource selection by sympatric free-ranging dairy cattle and brown bears Ursus arctos | 2011, Wildlife Biology: Volum 17 s. 389-403 Steyaert, Sam M.J.G; Støen, Ole-Gunnar; Elfström, Marcus; Karlsson, Jens; Van Lammeren, Ron; Bokdam, Jan
2022
Free-ranging cattle and bears in Sweden: are they compatible? 2022, Carnivore damage prevention news: s. 11-19 Swenson, Jon; Steyaert, Sam M.J.G.; Johnsen, Christin Beate; Rosell, Frank Narve; Zedrosser, Andreas
2021
Correction to: Size is not everything: differing activity and foraging patterns between the sexes in a monomorphic mammal (Behavioral Ecology and Sociobiology, (2021), 75, 4, (76), 10.1007/s00265-021-03010-7) 2021, Behavioral Ecology and Sociobiology: Volum 75 Lodberg-Holm, Hanna Kavli; Steyaert, Sam; Reinhardt, Stefanie; Rosell, Frank Narve
2019
Change in arthropod communities after a mass death of reindeer 2019, Birkemoe, Tone; Granum, Heidi Mørkhagen; Westermann, Lina; Aak, Anders; Frank, Shane; Steyaert, Sam
Change in arthropod communities following a mass death incident of reindeer at Hardangervidda
2019, Granum, Heidi Mørkhagen; Birkemoe, Tone; Aak, Anders; Steyaert, Sam
2012
How do 3,300 brown bears cope with 9.4 million Swedes? 2012, Swenson, Jon; Ordiz, Andres; Martin, Jodie; Steyaert, Sam M.J.G; Sahlén, Veronica; Støen, Ole-Gunnar